The removal of the linker histone should relax the constraints of the angle. The simulation is the same as above, except that the number of turns of DNA around the octamer is allowed to vary randomly between 1 and 2.
Yang, G., Leuba, S. H., Bustamante, C., Zlatanova, J. and van Holde, K. Role of linker histones in extended chromatin fibre structure. Nat Struct Biol. 1(11) 761-763 1994
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